Classification

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A tree is a plant form that occurs in many different orders and families of plants. Most species of trees today are flowering plants (angiosperms) and conifers. Trees show a variety of growth forms, leaf type and shape, bark characteristics and reproductive organs. For the listing of examples of well-known trees and how they are classified, see List of tree genera.

The tree form has evolved separately in unrelated classes of plants, in response to similar environmental challenges, making it a classic example of parallel evolution. With an estimate of 100,000 tree species, the number of tree species worldwide might total 25 percent of all living plant species. The majority of tree species grow in tropical regions of the world and many of these areas have not been surveyed yet by botanists, making species diversity and ranges poorly understood. The earliest tree-like organisms were tree ferns, horsetails and lycophytes, which grew in forests in the Carboniferous period, however these plants were not trees, since they lacked woody tissue. Trees evolved in the Triassic period, with conifers, ginkgos, cycads and other gymnosperms appeared producing woody tissue, and were subsequently followed by tree-form flowering plants in the Cretaceous period.

A small group of trees growing together is called a grove or copse, and a landscape covered by a dense growth of trees is called a forest. Several biotopes are defined largely by the trees that inhabit them; examples are rainforest and taiga (see ecozones). A landscape of trees scattered or spaced across grassland (usually grazed or burned over periodically) is called a savanna. A forest of great age is called old growth forest or ancient woodland (in the UK). A young tree is called a sapling.

Morphology

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The parts of a tree are the roots, trunk(s), branches, twigs and leaves. Tree stems consist mainly of support and transport tissues (xylem and phloem). Wood consists of xylem cells, and bark is made of phloem and other tissues external to the vascular cambium. Trees may be grouped into exogenous and endogenous trees according to the way in which their stem diameter increases. Exogenous trees, which comprise the great majority of trees (all conifers, and almost all broadleaf trees), grow by the addition of new wood outwards, immediately under the bark. Endogenous trees, mainly in the monocotyledons (e.g., aloes and dragon trees), grow by addition of new material as discrete bundles within the existing trunk tissue.

Trees can be identified to genus or species by a combination of the tree's shape, and the characteristics of its bark, leaves, flowers, and fruit. The leaves may be either deciduous or evergreen.

As an exogenous tree grows, it creates growth rings as new wood is laid down concentrically over the old wood. In species growing in areas with seasonal climate changes, wood growth produced at different times of the year may be visible as alternating light and dark, or soft and hard, rings of wood. In temperate climates, and tropical climates with a single wet-dry season alternation, the growth rings are annual, each pair of light and dark rings being one year of growth; these are known as annual rings. In areas with two wet and dry seasons each year, there may be two pairs of light and dark rings each year; and in some (mainly semi-desert regions with irregular rainfall), there may be a new growth ring with each rainfall. In tropical rainforest regions, with constant year-round climate, growth is continuous and the growth rings are not visible nor is there a change in the wood texture. In species with annual rings, these rings can be counted to determine the age of the tree, and used to date cores or even wood taken from trees in the past, a practice known as the science of dendrochronology. Very few tropical trees can be accurately dated in this manner. Age determination in this manner is also impossible in endogenous trees.

The roots of a tree are generally embedded in earth, providing anchorage for the above-ground biomass and absorbing water and nutrients from the soil. However, while ground nutrients are essential to a tree's growth the majority of its biomass – over 90 percent – comes from carbon dioxide absorbed from the atmosphere (see photosynthesis). Above ground, the trunk gives height to the leaf-bearing branches, aiding in competition with other plant species for sunlight. In many trees, the arrangement of the branches optimizes exposure of the leaves to sunlight.

Not all trees have all the plant organs or parts mentioned above. For example, the saguaro cactus of North America has no functional leaves. A plant form that is similar to a tree, but generally having smaller, multiple trunks and/or branches that arise near the ground, is called a shrub. However, no precise differentiation between shrubs and trees is possible. Given their small size, bonsai plants would not technically be "trees", but one should not confuse reference to the form of a species with the size or shape of individual specimens. A spruce seedling does not fit the definition of a tree, but all spruces are trees.